Forensic entomologist Dr John Manlove said it was his opinion that Mr Ryan’s body had remained in the tank “for most if not all of the time” since he went missing in June 2011.
Patrick Quirke (50), of Breanshamore, Co Tipperary, has pleaded not guilty to the murder of Mr Ryan (52) on a date between June 3, 2011, and April 2013.
Dr Manlove, who is based in the UK, explained to the jury that he specialises in the study of insects and certain invertebrates to help in investigations, particularly regarding suspicious deaths.
He told the trial at the Central Criminal Court that the body had not been exposed to insect activity for a long time.
He was engaged by An Garda Síochána to work on the case in 2014, he said. Specifically, he was asked whether an insect infestation had commenced in 2011 or in 2013.
Dr Manlove had access to evidence at the
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